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Termites can distinguish nestmates and non-nestmates through compound communication and gut symbionts: compounds consisting of hydrocarbons released from the cuticle permit the recognition of alien termite species.148149 Every colony has its own distinct odour. This odour is due to genetic and environmental elements like the termites' diet and the composition of these bacteria within the termites' intestines.150.

Termites rely on alarm communication to defend a colony.134 Alarm pheromones can be released when the nest has been breached or is being attacked by enemies or potential pathogens. Termites always avoid nestmates infected with Metarhizium anisopliae spores, through vibrational signals released by infected nestmates.151 Other procedures of defence include intense jerking and secretion of fluids in the frontal gland and defecating faeces containing alarm pheromones.134152.

 

 

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In some species, some soldiers obstruct tunnels to prevent their enemies from entering the nest, and they may intentionally repaint themselves as an act of defence.153 In scenarios where the intrusion is coming from a violation that is bigger than the soldier's head, defence requires a special formations where soldiers shape a phalanx-like formation around the breach and sting at intruders.154 If an invasion completed by Megaponera analis is successful, an entire colony may be destroyed, although this scenario is infrequent.154.

To termites, any breach of the tunnels or nests is a cause for alarm. When termites discover a potential violation, the soldiers usually bang their heads, apparently to attract other soldiers for defence and also to recruit additional employees to fix any breach.56 Additionally, an alarmed termite lumps into other termites which induces them to be alerted and to leave pheromone trails to the disturbed area, which can be a means to recruit additional workers.56.

 

 

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The pantropical subfamily Nasutitermitinae has a specialised caste of soldiers, known as nasutes, that have the capability to exude noxious fluids via a horn-like frontal projection they use for defence.155 Nasutes have lost their mandibles throughout the course of evolution and must be fed by workers.59 A wide variety of monoterpene hydrocarbon solvents have been identified in the liquids that nasutes secrete.156 Likewise, Formosan subterranean termites have been known to secrete naphthalene to safeguard their nests.157. over here

 

 

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Soldiers of those species Globitermes sulphureus commit suicide by autothysis  rupturing a large gland just beneath the surface of their cuticles. The thick, yellowish fluid in the gland becomes very tacky on contact with the atmosphere, entangling ants or other insects which are trying to invade the nest.158159 Another termite, Neocapriterme taracua, also engages in suicidal defence.

When soldiers guarding nest entrances are attacked by intruders, they engage in autothysis, creating a block that denies entry to any outsider.161.

Employees use several different strategies to deal with their dead, including burying, cannibalism, and avoiding a corpse altogether.162163164 To prevent pathogens, termites occasionally engage in necrophoresis, in which a nestmate carries a corpse in look at here now the colony to eliminate it elsewhere.165 Which strategy is used depends on the nature of this corpse a worker is dealing with (i.e.

 

 

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A species of fungus is known to mimic termite eggs, successfully avoiding its natural predators. These tiny brown chunks, known as"termite balls", seldom kill the eggs, and in some cases the workers tend to these.166 This fungus mimics these eggs by you can look here producing a cellulose-digesting enzyme known as glucosidases.167 A unique mimicking behavior exists between various species of Trichopsenius beetles and certain particular species within Reticulitermes.

This compound mimicry makes it possible for the beetles to integrate themselves within the termite colonies.168 The developed appendages on the physogastric abdomen of Austrospirachtha mimetes allows the beetle to mimic a termite worker.169.

Several species of ant are known to capture termites to utilize as a brand new food source later on, rather than killing them. By way of example, Formica nigra captures termites, and those who attempt to escape are immediately captured and driven underground.170 Certain species of ants in the subfamily Ponerinae conduct these raids although other ant species go in alone to steal the eggs or nymphs.146 Ants like Megaponera analis attack the outside of mounds and Dorylinae ants assault underground.146171 Despite this, some termites and ants can coexist peacefully.

 

 

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54 species of ants are known to inhabit Nasutitermes mounds, both occupied and abandoned ones.174 One reason many ants reside in Nasutitermes mounds is a result of the termites' frequent occurrence in their geographic range; another would be to protect themselves from flooding.174175 Iridomyrmex also inhabits termite mounds but no evidence for any kind of relationship (other than a one) is known.116 In rare cases, certain species of termites reside inside active ant colonies.176 Some invertebrate organisms such as beetles, caterpillars, flies and millipedes are termitophiles and dwell inside termite colonies (they are unable to survive independently).56 As a consequence, certain beetles and flies have evolved with their hosts.

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